Four acts performed on the Coachella Stage after the sun went down Friday, and Yeah Yeah Yeahs were the only ones who started releasing albums this century. Perhaps as a result, they couldn't play it safe, and they didn't. The band opened with their latest single, "Sacrilege" (performed again with a choir, this one imported from Los Angeles), and tested out the delicate, hushed new Mosquito track "Subway" directly after the crowd-pleasing hit "Zero." The taut punk of "Pin" and soupy dub of "Under the Earth" challenged the patience of chilled-out bros who seemed puzzled by the brilliant show-woman in a lavender jumpsuit cavorting in front of them. Karen O didn't double down on wild gestures to fill the massive stage, though, relying instead on a more subtle visual presentation paired with the band's superb musicianship. Karen managed to rouse the crowd from its stupor for a "Gold Lion" clap-along and the Show Your Bones gem came off full, bright, and inspired. Mosquito's "Slave" — performed for the second time ever, following its debut at New York's Webster Hall last weekend — was assured and haunting. And as designer Christian Joy correctly noted in our recent cover story, the band's most popular track these days appears to be "Heads Will Roll," which capped the 11-song set. It's hard to imagine the Strokes or any of the other bands birthed around the same time as the YYYs stepping onto such a massive platform and throwing down the gauntlet like this, challenging the audience to get with a brand-new program. Which is probably why Yeah Yeah Yeahs are the only band of its generation with such plumb billing this year.